However, human or not, a soft susurrus of language washed over the group. The talking was among the mothers and infants, the groomers, and the couples. There wasn’t much information being passed on; much of the talk was little more than sighs of pleasure, like the purring of cats.

But their words sounded like words.

People had had to learn to communicate with equipment designed for other tasks — a mouth intended for eating, ears intended to listen for danger — now jury-rigged for a new use. Their bipedalism had helped: the repositioning of their larynxes and changes in the pattern of breathing improved the quality of the sounds they could make. But to be useful, sounds had to be identifiable quickly and unambiguously. And the ways the hominids could achieve that were limited by the nature of the equipment they had to use. As people listened to each other, and imitated and reused useful noises, phonemes — the sound content of the words, the basis of all language — had selected themselves, driven by communicative necessity and engineering limitations.

But there was nothing yet like grammar — no sentences — and certainly no narratives, no stories. And the main purpose of talking right now wasn’t to pass on information. Nobody talked about tools or hunting or food preparation. Language was social: It was used for commands and demands, for blunt expressions of joy or pain. And it was used for grooming: Language, even without much content, was a more efficient way to establish and reinforce relations than picking ticks out of pubic hair. It even worked to “groom” several people at once.

A lot of the evolution of language, in fact, had been driven by mothers and infants. Right now the ancestors of Demosthenes and Lincoln and Churchill spoke nothing much more than motherese.

And the children didn’t talk at all.

The minds of the adults were about equivalent in complexity to a five-year-old human’s. Their children were not capable of speech — nothing beyond chimplike jabbers — until they reached adolescence. It had only been a year or two since the adults’ words had made any sense to Far, and the Brat, at seven, couldn’t talk at all. The kids were like apes born to human parents.

As the light died, so the group settled toward sleep.

Far huddled against her mother’s legs. The ending day became just one of a long chain that stretched back to the beginning of her life, days dimly remembered, only vaguely linked. In the darkness she imagined running in the blinding brightness of day, running and running.

She had no way of knowing that this was the last time she would fall asleep close to her mother.

A million years ago, tectonic drift, slow but relentless, had caused North and South America to collide, and the isthmus of Panama was formed.

In itself it seemed a small event, Panama an inconsequential sliver of land. But, as with Chicxulub, this region had once more become the epicenter of a worldwide catastrophe.

Because of Panama, the old equatorial flows through the Americas — the last trace of the Edenic Tethys current — had been cut off. Now the only Atlantic currents were the huge interpolar flows, great conveyor belts of cold water. The worldwide cooling intensified drastically. The scattered ice caps covering the northern ocean merged, and glaciers spread like claws over the northern landmasses.

The Ice Ages had begun. At their greatest extent the glaciers would cover more than a quarter of all Earth’s surface; the ice would reach as far as Missouri and central England. Much was immediately lost. Where the glaciers passed, the land was scraped clean — down to the bedrock, which was itself pulverized and ground to dust — leaving a legacy of mountains with scored flanks, polished surfaces, scattered boulders, and gouged-out valleys. There had been no significant glaciation on Earth for two hundred million years; now a legacy of rocks and bones dating back deep into the age of the dinosaurs was comprehensively destroyed.

On the ice itself, nothing could live: nothing. Below the ice, great impoverished belts of tundra spread. Even in places far from the ice, like the equatorial regions of Africa, changes in wind patterns intensified the aridity, and vegetation shrank back to the coasts and river valleys.

The cooling was not a uniform trend. The planet tipped and bobbed in its endless dance around the sun, subtly shifting its degree of tilt, its inclination, and the fine-tuning of its orbit. And with each cycle the ice came and went, came and went; ocean levels fluctuated like the pumping of a heart. Even the land, compressed under kilometers of ice or released by its melting, rose and fell like a rocky tide.

Sometimes the climate shifts could be savage. Within a single year the amount of snowfall in an area could double, the average temperature fall by ten degrees. Faced with such chaotic oscillations, living things moved, or died.

Even the forests marched. Spruce proved a fast migrant, followed by pine, capable of marching at a kilometer every two years. The great chestnuts, massive trees with heavy seeds, could manage a pace of a hundred meters a year. Before the Ice Age the animals of the middle latitudes of the northern hemisphere had been a rich mixture of fleet grazers like deer and horses, giant herbivores like rhinos, and fast-running carnivores like lions and wolves. Now the animals were driven south in search of warmth. Populations of animals from different climatic zones were mixed up and forced to compete in fast-changing ecological arenas.

But some creatures began to adapt to the cold, to exploit the food supplies that still existed at the feet of the ice sheets. Many animals grew thick fur and layers of fat — large animals, like rhinos, and smaller animals, like foxes and horses and cats. Others began to take advantage of the huge temperature swings between the seasons. They migrated, moving north in the summer and south in the winter; the plains became a huge tidal wash of life, great mobile communities patiently stalked by predators.

There had been a catastrophe of mixing in the Americas. The two continents, north and south, had been separated since the shattering of Pangaea some one hundred and fifty million years ago. The fauna of South America had evolved in isolation, and was dominated by marsupial mammals and ungulates. There were marsupial “wolves” and saber-toothed “cats”; there were ungulate “camels” and trunked “elephants,” and giant ground sloths that could weigh three tons and stood six meters tall when they stood up to browse on palm leaves. There were still glyptodonts, not so dissimilar from the huge armored beast that had terrified Roamer, and the top predators were giant flightless birds, just as in archaic times. This exotic assemblage had been left alone to develop — though it was supplemented from time to time by waifs, brought by rafting or temporary bridges, like Roamer and her hapless companions, whose children had populated the South American jungles with monkeys.

But when the Panama land bridge was closed there was a massive migration from north to south of insectivores, rabbits, squirrels, mice, and later dogs, bears, weasels, and cats. The natives of South America failed to compete with these new arrivals. The extinctions took millions of years, but the empire of the marsupials was done.

For all the difficulty and dying, this time of fast and savage changes was, perversely, a time of opportunity. In the entire four-billion-year history of the Earth there had been few times more propitious for diversification and evolutionary innovation. Amid much extinction there was wild speciation.

And right at the center of this ecological cauldron were the children of Capo.

The next morning dawned brightly, with a washed-out blue sky. But the air was very dry and smelled oddly sharp, and the heat was soon stifling. The animals of the savannah seemed subdued. Even the birds were quiet; the carrion eaters clung to their tree roosts like ugly black fruit.